Sarah Gergel, Professor, Landscape Ecology & Conservation

Landscape Ecology Lab, Forest and Conservation Sciences

2nd Edition

We have been extremely pleased and honored by the enthusiasm generated by the first edition of Learning Landscape Ecology, the first “hands-on” lab textbook for learning the concepts and tools of the discipline. We’re excited that the text has been incorporated into the curricula of more than 55 universities world-wide, and has even been translated into Korean.

The second edition is decidedly more applied and international in the choice of examples, approaches, perspectives, and contributors. It includes new advances in quantifying landscape structure and connectivity (such as graph theory), as well as labs which incorporate the latest scientific understanding of ecosystem services, resilience, social-ecological landscapes, and even seascapes.

As in the 1st edition, the exercises emphasize easy-to-use, widely-available software. Introductory exposure to spatial analyses using R programming language is provided by several labs. What remains similar from the 1st edition is our dedication to making seemingly complex ideas easy to understand and use for scientists from diverse backgrounds, and particularly for those early in their careers.

All the files, software and datasets are available using the links below for a given chapter. For a complete copy of the Instructor’s Notes (with Answer Keys), instructors can email SEG: with your course number, title and affiliation.

Sarah Gergel and Monica Turner, Editors


Some links have changed (as of Sept 2021), so be sure to refresh your browser a few times if you are being denied access to any folders.

Advice for Instructors


Chapter 1: Introduction to Remote Sensing 
Nicholas C. Coops and T. Rory Tooke

Chapter 2: Historical Aerial Photographs for Landscape Analysis
Jessica L. Morgan, Sarah E. Gergel, Collin Ankerson, Stephanie A. Tomscha and Ira Sutherland

Chapter 3: Citizen Science for Assessing Landscape Change
Jeffrey A. Cardille and Michelle M. Jackson


Chapter 4: Understanding Landscape Metrics
Jeffrey A. Cardille and Monica G. Turner

Chapter 5: Scale Detection with Semivariograms and Autocorrelograms (with R option)
Michael W. Palmer and Daniel J. McGlinn

Chapter 6: Characterizing Categorical Map Patterns Using Neutral Landscape Models (with QRULE and R)
Robert H. Gardner

Chapter 7: What Constitutes a Significant Difference in Landscape Pattern? (using R)
Tarmo K. Remmel and Marie-Josée Fortin


Chapter 8: Modeling Landscape Change with Markov Models (with R option)
Dean L. Urban and David O. Wallin

Chapter 9: Simulating Management Actions and Their Effect on Forest Landscape Pattern (with Harvest Lite)
Eric J. Gustafson

Chapter 10: Regional and Continental-scale Perspectives on Landscape Pattern
Jeffrey A. Cardille and Monica G. Turner

Chapter 11: Using Spatial Statistics and Landscape Metrics to Compare Disturbance Mosaics (with GS+)
Monica G. Turner and Martin Simard


Chapter 12: Assessing Multi-scale Landscape Connectivity Using Network Analysis  Todd R. Lookingbill and Emily S. Minor

Chapter 13: Conservation Planning (with Marxan)
Matthew Watts, Hugh P. Possingham, Carissa J. Klein, Tara G. Martin and Josie Carwardine

Chapter 14: Advances in Quantifying Habitat Connectivity Using Graph Theory (with Conefor)
Santiago Saura and Begoña de la Fuente

Chapter 15: Linking Landscapes & Meta-Communities (using R)
Ben Gilbert and Joseph R. Bennett


Chapter 16: Modelling Spatial Dynamics of Ecosystem Services
Sarah E. Gergel and Tara Reed

Chapter 17: Heterogeneity in Ecosystem Services: Multi-scale Carbon Management in Tropical Forest Landscapes
Kathryn R. Kirby, Jeanine M. Rhemtulla and Sarah E. Gergel

Chapter 18: Regime Shifts and Spatial Resilience in a Coral Reef Seascape
Jennifer C. Selgrath, Garry D. Peterson, Matilda Thyresson, Magnus Nyström and Sarah E. Gergel

Chapter 19: Understanding Land-Use Feedbacks and Ecosystem Service Tradeoffs in Agriculture
Lisa A. Schulte and John C. Tyndall

Chapter 20: Social Networks: Uncovering Social-ecological Mismatches in Heterogeneous Marine Landscapes
Örjan Bodin and Beatrice I. Crona


Files from the first edition (2001) 


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Forest and Conservation Sciences
Forest Sciences Centre
2424 Main Mall,
Vancouver, BC V6T 1Z4, Canada

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